Volcano

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Hugo and Basillus are the black plauge representing the Justinian plague and black death respectively. Astrion is the Thera Volcano the pushed the Minoan civilization in to decline. This difference is why we see different abilities with the threes manifestation of the macula. Asterion turns people to ash statues like in Pompeii (something some archeologists believed haped to people during Thera Eruption) this is also why his powers involve tentacles that Erupted from the earth that turn in to ash. Astrion and his macula might also only be genetically passed by the people of his island wich means it might be extinct after his death. Also Astrion is the actual name of the minotaur and his mother most likely died in childbirth.